Aftereffect of BMPWntNodal transmission upon originate cellular distinction

In those groups, signing is used for hunting and for a rich array of ritual purposes, including mourning and traditional story-telling. We discuss further evidence, based on typological generalizations about the phonology of non-primary sign languages and comparative ethnographic work, that points to such a parallelism. This evidence includes the fact that for some of those groups, stencil and petroglyph art has independently been linked to their sign language expressions. This article is part of the theme issue 'Reconstructing prehistoric languages'.Bodily mimesis, the capacity to use the body representationally, was one of the key innovations that allowed early humans to go beyond the 'baseline' of generalized ape communication and cognition. We argue that the original human-specific communication afforded by bodily mimesis was based on signs that involve three entities an expression that represents an object (i.e. communicated content) for an interpreter. We further propose that the core component of this communication, pantomime, was able to transmit referential information that was not limited to select semantic domains or the 'here-and-now', by means of motivated-most importantly iconic-signs. Pressures for expressivity and economy then led to conventionalization of signs and a growth of linguistic characteristics semiotic systematicity and combinatorial expression. Despite these developments, both naturalistic and experimental data suggest that the system of pantomime did not disappear and is actively used by modern humans. Its contemporary manifestations, or pantomimic fossils, emerge when language cannot be used, for instance when people do not share a common language, or in situations where the use of (spoken) language is difficult, impossible or forbidden. Under such circumstances, people bootstrap communication by means of pantomime and, when these circumstances persist, newly emergent pantomimic communication becomes increasingly language-like. This article is part of the theme issue 'Reconstructing prehistoric languages'.Two families of quantitative methods have been used to infer geographical homelands of language families Bayesian phylogeography and the 'diversity method'. Bayesian methods model how populations may have moved using a phylogenetic tree as a backbone, while the diversity method assumes that the geographical area where linguistic diversity is highest likely corresponds to the homeland. No systematic tests of the performances of the different methods in a linguistic context have so far been published. Here, we carry out performance testing by simulating language families, including branching structures and word lists, along with speaker populations moving in space. We test six different methods two versions of BayesTraits; the relaxed random walk model of BEAST 2; our own RevBayes implementations of a fixed rate and a variable rates random walk model; and the diversity method. As a result of the tests, we propose a hierarchy of performance of the different methods. Factors such as geographical idiosyncrasies, incomplete sampling, tree imbalance and small family sizes all have a negative impact on performance, but mostly across the board, the performance hierarchy generally being impervious to such factors. This article is part of the theme issue 'Reconstructing prehistoric languages'.In this paper, we compare the languages each of the authors invented as prehistoric languages for popular culture media. Schreyer's language, Beama, was created for the film Alpha (2018), while Adger's language, Tan!aa Kawawa ki, was created for a television series on how early hominins spread throughout the world (the series was green-lit but then cancelled). We argue that though this creative process may seem far removed from classical research paradigms on language evolution, it can provide some insight into how disparate research on the possible properties of prehistoric languages can be brought together to illustrate how these languages might have worked as whole linguistic systems within these imagined worlds, as well as in prehistory. Eliglustat This article is part of the theme issue 'Reconstructing prehistoric languages'.While speech and language do not fossilize, they still leave traces that can be extracted and interpreted. Here, we suggest that the shape of the hard structures of the vocal tract may also allow inferences about the speech of long-gone humans. These build on recent experimental and modelling studies, showing that there is extensive variation between individuals in the precise shape of the vocal tract, and that this variation affects speech and language. In particular, we show that detailed anatomical information concerning two components of the vocal tract (the lower jaw and the hard palate) can be extracted and digitized from the osteological remains of three historical populations from The Netherlands, and can be used to conduct three-dimensional biomechanical simulations of vowel production. We could recover the signatures of inter-individual variation between these vowels, in acoustics and articulation. While 'proof-of-concept', this study suggests that older and less well-preserved remains could be used to draw inferences about historic and prehistoric languages. Moreover, it forces us to clarify the meaning and use of the uniformitarian principle in linguistics, and to consider the wider context of language use, including the anatomy, physiology and cognition of the speakers. This article is part of the theme issue 'Reconstructing prehistoric languages'.The aim of this paper is to develop further the idea that symptoms that emerge in speech and language processing following brain damage can make a contribution to discussions of the early evolution of language. These diverse impairments are called aphasia, and this paper proposes that the recovery of a non-fluent aphasia syndrome following stroke could provide insights into the course of the pre-history of human language evolution. The observable symptoms emerge during recovery, crucially enabled by (dis)inhibition in parallel with a range of impairments in action processing (apraxias), including apraxia of speech. They are underpinned by changes in cortical and subcortical status following brain damage. It is proposed that the observed recovery mimics ontogenic and phylogenic processes in human speech and language. The arguments put forward provide insights tending to support the motor-gestural model of speech and language evolution. This article is part of the theme issue 'Reconstructing prehistoric languages'.